BACKGROUND CONCEPTS

1. Our eyes allow us to perceive electromagnetic radiation reflected from objects

Most animals and many plants are photosensitive; that is, they can detect different light intensities. Some organisms accomplish this with single cells or with simple eyes that do not form images but do allow the organism to react to light by moving toward or away from it. In order for an eye to transmit more information about the world, however, it must have a way of forming an image, a representation of the scene being viewed.

Higher invertebrates and virtually all vertebrates have complex, image-forming eyes, and we will "focus" on the refracting eye found in the octopus and in all vertebrates. Arthropods have compound eyes, which have greater depth of focus than refracting eyes, but which sacrifice resolving power or acuity. Our eyes, like those of many animals, detect a just narrow range of all the wavelengths of electromagnetic radiation, that between 380 and 760 nanometers. This range of light is called the visible spectrum. Figure 1 shows how the visible spectrum fits into the entire electromagnetic spectrum.

Figure 1. The electromagnetic spectrum and the visible spectrum.

2. The eyeball is an optical device for focusing light

The mammalian eyeball (Figure 2) is an organ that focuses a visual scene onto a sheet of specialized neural tissue, the retina, which lines the back of the eye. Light from a scene passes through the cornea, pupil, and lens on its way to the retina. The cornea and lens focus light from objects onto photoreceptors, which absorb and then convert it into electrical signals that carry information to the brain. Two pockets of transparent fluid nourish eye tissues and maintain constant eye shape: these are the aqueous and vitreous humors, through which the light also passes. The lens projects an inverted image onto the retina in the same way a camera lens projects an inverted image onto film; the brain adjusts this inversion so we see the world in its correct orientation. To control the images that fall upon our retinas, we can either turn our heads or turn our eyes independently of our heads by contracting the extraocular muscles, six bands of muscles that attach to the tough outside covering, or sclera, of the eyeball and are innervated by cranial nerves. See Table 1 for a brief list of eyeball components and their functions.

The cornea and lens bend or refract light rays as they enter the eye, in order to focus images on the retina. The eye can change the extent to which rays are bent and thus can focus images of objects that are various distances from the observer, by varying the curvature of the lens. The ciliary muscle accomplishes this by contracting to lessen tension on the lens and allowing it to round up so it can bend light rays more, or relaxing for the opposite effect. This ciliary muscle is smooth or non-voluntary muscle-you cannot "decide" to contract or relax it as you do the skeletal muscle in a finger or facial muscle.

Figure 2. The mammalian eyeball.

3. Refractive errors in the eye cause focusing problems

Refractive errors occur when the bending of light rays by the cornea and lens does not focus the image correctly onto the retina. An eyeball that is too long or too short for the optics of the cornea and lens or an irregularly shaped cornea can cause refractive errors, which include myopia (near-sightedness), hyperopia (far-sightedness), and astigmatism. Myopia results either when the eyeball is too long or when the cornea is curved too much, and the focused image falls in front of the retina. Hyperopia is the opposite, with the image falling behind the retina. Astigmatism results from a cornea that is not spherical. Fortunately, most refractive errors can be corrected with prescription lenses.